The four consequences, in the plans' own words. First, the minor owns it. my529's Form 104 states that "for an UGMA/UTMA account, the minor is both the account owner and beneficiary". Second, the beneficiary is fixed: "the UGMA/UTMA account agent cannot change the beneficiary on this account or make account withdrawals other than for the benefit of the beneficiary", and Invest529 lists "inability to change the Beneficiary" among its custodial-account restrictions. Third, no successor owner may be named, because "the account money is a permanent gift to the beneficiary"; Invest529 reaches the same place from the other direction by requiring that "custodial Accounts ... must name the Beneficiary's estate as the Designated Survivor". Fourth, and following from the third, "if the beneficiary of an UGMA/UTMA account dies, the account money will become part of the estate of that beneficiary" rather than reverting to whoever funded it.
None of those is an ownership label on an ordinary account. Each is a restriction on what can be done with the money, which is why the plans administer it as its own account type and require it to be "established separately from any other accounts that the UGMA/UTMA account agent may hold for the beneficiary".
The person managing it has a different title and a different job. my529 calls them the "UGMA/UTMA account agent" rather than a custodian, notes that the agent "is not required to be the custodian of any former UGMA/UTMA account", and ties the end of their authority to "the age of majority in the state in which the money was originally gifted", not the state of the plan. At that point the agent files a change form and, in my529's words, "the account type will be changed from UGMA/UTMA to individual". Invest529 is explicit that nothing happens automatically: it "will not automatically transfer the UTMA/UGMA account to the Beneficiary when they reach the age of majority" and must be notified with documentation that the custodianship has terminated.
The conversion is one-directional, and it contaminates in one direction too. Existing custodial money must be liquidated before it can be contributed, since a 529 takes cash rather than securities, and my529 warns that a custodian "should discuss any potential tax consequences of liquidating an UGMA/UTMA account with their tax advisors" first, because the sale is a taxable event in the child's hands. Going the other way, my529 states that "once contributions are made to an UGMA/UTMA account, the entire account is subject to custodial account and UGMA/UTMA rules, regardless of the source of the contributions", and Invest529 that "non-custodial funds contributed to the custodial account are irrevocable gifts to the minor and become custodial funds". So adding ordinary family money to a custodial 529 converts that money into the child's property. Anyone wanting flexible money for the same child opens a second, ordinary account.
The federal aid treatment is why families do this at all, and it is counter-intuitive. Under 20 U.S.C. 1087vv(f)(3), a qualified education benefit "shall be considered an asset of ... the parent if the student is a dependent student and the account is designated for the student, regardless of whether the owner of the account is the student or the parent", and subsection (f)(4)(A) defines qualified education benefit to include a 529 program. So a 529 owned by a dependent student is reported on the federal aid application as a parent asset. Parents' assets are reduced by an age-based asset protection allowance and then counted at 12 percent under 20 U.S.C. 1087oo(d)(1)(A) before entering the parents' assessment schedule, while 1087oo(h) counts a student's own assets at a flat 20 percent with no allowance. A plain custodial brokerage account is the student's own property and is counted the harder way. Moving that money into a custodial 529 keeps every custodial restriction and changes which line of the aid formula it lands on. Invest529's program description states the federal rule in the same terms, adding that institutional formulas may differ.
Two tax points to keep separate. The 529's own federal tax treatment does not change because the account is custodial: growth is tax-deferred and qualified distributions are federally tax-free either way. What does change is who owns the tax attributes at state level. Invest529's rule is that the Virginia deduction "for UTMA/UGMA Invest529 Accounts belongs to the Beneficiary, and is reported under the Beneficiary's Social Security number", and that custodians are not eligible for it on their contributions. Whether a state offers a deduction or credit, and to whom, is a state-by-state question, and it is one of the few places where a custodial 529 is worse than an ordinary one for the adult writing the checks.